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No, Part of the definition of ProneDDI is: You can’t move from your space, although you can teleport, crawl, or be forced to move by a pull, a push, or a slide. Lightning shift does not use one of the forms of movement that can allow a creature to move while prone.

Per the rules compendium page 249, you can't shift over any terrain that requires a skill check. So unless you have a swim speed, you can't shift while in water. Here's the excerpt from the Rules Compendium - see the Special Movement Modes section: Shift Action: Move action. Movement: The creature moves 1 square. (Some powers and effects allow ...

You can shift while swimming, but remember a few things about aquatic movement: You must make an athletics check to swim. You must spend 1 extra square of movement for each square you enter while swimming. Full rules, including DC's for different types of water, can be found under the athletics skill. Therefore, the normal move action of shifting 1 ...

In Pathfinder, natural attacks are treated somewhat differently from weapon attacks. There is also a section on natural attacks under the Attack Action (not the Full Attack Action) in the Combat section. But the Combat section clearly says, after talking about all these kinds of attacks (weapon, unarmed, natural) that "A character who can make more than one ...

In order to use Lightning Shift while prone, you need to take one of a number of feats or items: In rough order from least onerous to most: Feet slot: Floorfighter Straps Feat: Low Crawl Acrobat's Boots and other effects: while using a minor to stand up isn't exactly shifting while prone, they allow you to shift on the same turn that you were prone. ...