This question got me thinking about ways to let a druid wildshaped into a giant squid walk or fly permanently.
While searching for ways I stumbled upon the synergist witch.
It can merge with her familiar using her symbiosis ability and gets some of her familiar's abilities while doing so. One of them is:
At 11th level, during symbiosis the synergist gains any abilities the familiar possesses that are listed under beast shape III. If the familiar can fly, the synergist can fly for an unlimited amount of time each day while in symbiosis.
Is there any reason for symbiosis to not work during wildshape? It is not an ability based on her normal form because it is an ability she gets from merging with her familiar.
Because of the high levels in both druid and witch this is probably something only relevant as a thought experiment. At first I thought 5 levels synergist were enough to get a walking speed untill I rechecked and had to notice that nothing in the form of a walking speed was listed under beast shape.